Template variables largely work in exact the same way as skin variables, with the only difference that they are being used inside templates. The variables are called using <%varname%>, and include some text depending on the variable function. Some variables also have extra optional parameters.
These template variables be used in the following template-parts: item header, item body, item footer, date header, date footer, morelink, editlink (The variables image, popup and media can also be used inside weblog items.)
Comments-related template-parts (comments header, comments body, comments footer, one comment, two comments, comments read more, no comments, too much comments) have a different set of available variables: